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Common name Gene Organism Uniprot Acc Name Organelle In vivo In vitro Membrane cluster Partner dependent RNA Dependency PTM Dependency Domain-motif interaction Discrete oligomerization
Whi3 AGOS_ABL158C Ashbya gossypii Q75E28 ABL158Cp
cytoplasmic stress granule
T
T
F
F
F
F
F
F
LAF-1 LAF-1 Caenorhabditis elegans D0PV95 ATP-dependent RNA helicase laf-1
P granule
T
T
F
F
F
F
F
F
BugZ ZNF207 Xenopus laevis Q7ZXV8 BUB3-interacting and GLEBS motif-containing protein ZNF207
spindle matrix
T
T
F
F
F
F
F
F
FUS FUS Homo sapiens P35637 RNA-binding protein FUS
cytoplasmic stress granule; cytoplasmic ribonucleoprotein granule
T
T
F
F
F
F
F
F
FMRP FMR1 Homo sapiens Q06787 Synaptic functional regulator FMR1
cytoplasmic stress granule; cytoplasmic ribonucleoprotein granule; synaptosome, neuron projection; neuronal ribonucleoprotein granule
T
T
F
T
T
F
F
F
TAF15 TAF15 Homo sapiens Q92804 TATA-binding protein-associated factor 2N
nuclear protein granule
T
T
F
F
F
F
F
F
TDP-43, TDP43 TARDBP Homo sapiens Q13148 TAR DNA-binding protein 43
cytoplasmic stress granule; cytoplasmic ribonucleoprotein granule
T
T
F
F
F
F
F
F
Ataxin-2 ATX2 Drosophila melanogaster Q8SWR8 Ataxin-2 homolog
neuronal ribonucleoprotein granule; cytoplasmic stress granule
T
T
F
F
T
F
F
F
EWS EWSR1 Homo sapiens Q01844 RNA-binding protein EWS
nuclear protein granule
T
T
F
F
F
F
F
F
HNRNPA1, HNRPA1 HNRNPA1 Homo sapiens P09651 Heterogeneous nuclear ribonucleoprotein A1
cytoplasmic stress granule
T
T
F
F
F
F
F
F
HNRNPA2B1, HNRPA2B1 HNRNPA2B1 Homo sapiens P22626-2 Isoform A2 of Heterogeneous nuclear ribonucleoproteins A2/B1
cytoplasmic stress granule; cytoplasmic ribonucleoprotein granule
T
T
F
F
F
F
T
F
TIA1 TIA1 Homo sapiens P31483 Nucleolysin TIA-1 isoform p40
cytoplasmic stress granule
T
T
F
F
F
F
F
T
Nephrin NPHS1 Homo sapiens O60500 Nephrin
membrane cluster; actin cortical patch; Arp2/3 protein complex
T
T
T
T
F
T
T
F
N-WASP WASL Homo sapiens O00401 Neural Wiskott-Aldrich syndrome protein
membrane cluster; actin cortical patch; Arp2/3 protein complex
T
T
T
T
F
T
T
F
NCK1, Nck NCK1 Homo sapiens P16333 Cytoplasmic protein NCK1
membrane cluster; actin cortical patch; Arp2/3 protein complex
T
T
T
T
F
T
T
F
DDX4 DDX4 Homo sapiens Q9NQI0 Probable ATP-dependent RNA helicase DDX4
P granule; pi-body
T
T
F
F
F
F
F
F
DDX3 DDX3X Homo sapiens O00571 ATP-dependent RNA helicase DDX3X
cytoplasmic stress granule
T
T
F
F
F
F
F
F
NUP98 NUP98 Homo sapiens P52948 Nuclear pore complex protein Nup98-Nup96
nuclear pore central transport channel; selective hydrogel-like meshwork formed by FG-nucleoporins in nuclear pore central channel
T
T
F
F
F
F
F
F
Nsp1, Nsp1p NSP1 Saccharomyces cerevisiae P14907 Nucleoporin NSP1
nuclear pore central transport channel; selective hydrogel-like meshwork formed by FG-nucleoporins in nuclear pore central channel
T
T
F
F
F
F
F
F
Nup100 NUP100 Saccharomyces cerevisiae Q02629 Nucleoporin NUP100/NSP100
nuclear pore central transport channel; selective hydrogel-like meshwork formed by FG-nucleoporins in nuclear pore central channel
T
T
F
F
F
F
F
F
Elastin ELN Homo sapiens P15502 Elastin
extracellular matrix; collagen-containing extracellular matrix
F
T
F
F
F
F
F
F
PGL-3 PGL-3 Caenorhabditis elegans G5EBV6 Guanyl-specific ribonuclease pgl-3
P granule
T
T
F
F
T
F
F
T
MEG-3 MEG-3 Caenorhabditis elegans Q9TXM1 Uncharacterized protein
P granule
T
T
F
F
F
F
F
F
Pub1 PUB1 Saccharomyces cerevisiae P32588 Nuclear and cytoplasmic polyadenylated RNA-binding protein PUB1
cytoplasmic stress granule
T
T
F
F
F
F
F
F
eIF4GII TIF4632 Saccharomyces cerevisiae P39936 Eukaryotic initiation factor 4F subunit p130
cytoplasmic stress granule
F
T
F
F
F
F
F
F
RBM14 RBM14 Homo sapiens Q96PK6 RNA-binding protein 14
paraspeckle
T
T
F
F
F
F
F
F
HP1α CBX5 Homo sapiens P45973 Chromobox protein homolog 5
heterochromatin
T
T
F
F
F
F
F
F
HP1a SU(VAR)205 Drosophila melanogaster P05205 Heterochromatin protein 1
heterochromatin
T
T
F
F
F
F
F
F
PAB1 PAB1 Saccharomyces cerevisiae P04147 Polyadenylate-binding protein, cytoplasmic and nuclear
cytoplasmic stress granule
T
T
F
F
F
F
F
F
Fibrillarin, fib-1 FIB-1 Caenorhabditis elegans Q22053 rRNA 2-O-methyltransferase fibrillarin
nucleolus; dense fibrillar component; nuclear body
T
T
F
T
T
F
F
F
NPM1 NPM1 Homo sapiens P06748 Nucleophosmin
nucleolus; granular component
T
T
F
T
T
F
T
T
SPD-5 SPD-5 Caenorhabditis elegans P91349 Spindle-defective protein 5
pericentriolar material
T
T
F
F
F
T
F
F
UBQLN2 UBQLN2 Homo sapiens Q9UHD9 Ubiquilin-2
cytoplasmic stress granule
T
T
F
F
F
F
T
F
cpeb4 CPEB4 Xenopus laevis D8V196 Cytoplasmic polyadenylation element binding protein 4
cytoplasmic protein granule
T
F
F
F
F
F
F
F
HSPB2 HSPB2 Homo sapiens Q16082 Heat shock protein beta-2
cytoplasmic protein granule; nuclear protein granule
T
F
F
F
F
F
F
F
Tau-F, Tau MAPT Homo sapiens P10636-8 Isoform Tau-F of Microtubule-associated protein tau
cytoplasmic microtubule; condensed compartments of microtubule bundling
T
T
F
F
F
F
F
F
TNRC6B, GW182 TNRC6B Homo sapiens Q9UPQ9 Trinucleotide repeat-containing gene 6B protein
RISC complex; P-body; micro-ribonucleoprotein complex; miRISC; GW-body
T
T
F
T
F
F
T
F
SynGAP SYNGAP1 Mus musculus J3QQ18 Ras/Rap GTPase-activating protein SynGAP
postsynaptic density
T
T
F
T
F
F
T
T
Galectin-3 LGALS3 Homo sapiens P17931 Galectin-3
galectin complex; galectin lattice
F
T
T
F
F
F
F
T
Rbfox1 RBFOX1 Drosophila melanogaster Q7YU08 LD15974p
cytoplasmic ribonucleoprotein granule; nuclear body
T
F
F
F
T
T
F
F
p62 SQSTM1 Homo sapiens Q13501 Sequestosome-1
inclusion body; p62 body
T
T
F
F
F
T
F
T
LAT LAT Homo sapiens O43561 Linker for activation of T-cells family member 1
TCR signalosome; LAT signalosome
T
T
T
T
F
T
T
F
GRB2 GRB2 Homo sapiens P62993 Growth factor receptor-bound protein 2
TCR signalosome; LAT signalosome
T
T
T
T
F
T
T
F
SOS1 SOS1 Homo sapiens Q07889 Son of sevenless homolog 1
TCR signalosome; LAT signalosome
T
T
T
T
F
T
T
F
NONO NONO Homo sapiens Q15233 Non-POU domain-containing octamer-binding protein
paraspeckle
T
F
F
T
T
T
F
T
SFPQ SFPQ Homo sapiens P23246 Splicing factor, proline- and glutamine-rich
paraspeckle
T
F
F
T
T
T
F
T
Fib FIB Drosophila melanogaster Q9W1V3 rRNA 2-O-methyltransferase fibrillarin
nucleolus; dense fibrillar component; nuclear body
T
F
F
F
F
T
T
T
Nopp140 NOPP140 Drosophila melanogaster Q7KTV5 Nopp140, isoform B
nucleolus
T
F
F
F
F
T
T
T
Pit PIT Drosophila melanogaster Q9VD51 Probable ATP-dependent RNA helicase pitchoune
nucleolus
T
F
F
F
F
T
T
T
Pbp1 PBP1 Saccharomyces cerevisiae P53297 PAB1-binding protein 1
cytoplasmic stress granule
T
T
F
F
F
F
F
T
Dcp2 DCP2 Schizosaccharomyces pombe O13828 mRNA decapping complex subunit 2
P-body
T
T
F
T
F
F
F
F
Edc3 EDC3 Schizosaccharomyces pombe O94752 Enhancer of mRNA-decapping protein 3
P-body
T
T
F
T
F
F
F
F
Pdc1 SPAC13A11.06 Schizosaccharomyces pombe Q09737 Putative pyruvate decarboxylase C13A11.06
P-body
T
T
F
T
F
F
F
F
Pon PON Drosophila melanogaster Q9W4I7 Partner of numb
cytoplasmic protein granule; basal Numb-Pon crescent in dividing neuroblasts
T
T
F
T
F
F
T
F
NUMB NUMB Drosophila melanogaster P16554 Protein numb
cytoplasmic protein granule; basal Numb-Pon crescent in dividing neuroblasts
T
T
F
T
F
F
T
F
Sup35 SUP35 Saccharomyces cerevisiae P05453 Eukaryotic peptide chain release factor GTP-binding subunit
cytoplasmic protein granule; Sup35 condensate
T
T
F
F
F
F
F
F
MUT-16 MUT-16 Caenorhabditis elegans O62011 MUTator
mutator focus; small RNA amplification complex
T
T
F
F
F
F
F
F
SYN1 SYN1 Homo sapiens P17600 Synapsin-1
presynaptic cytosol; a matrix holding together clusters of synaptic vesicles (SVs)
T
T
F
F
F
F
F
T
cGAS CGAS Homo sapiens Q8N884 Cyclic GMP-AMP synthase
intracellular DNA/protein granule; cGAS foci
T
T
F
T
F
F
F
T
MED1 MED1 Homo sapiens Q15648 Mediator of RNA polymerase II transcription subunit 1
enhanceosome; nuclear body; nuclear bodies that occur at super enhancers in mESCs
T
T
F
F
F
F
F
F
BRD4 BRD4 Homo sapiens O60885 Bromodomain-containing protein 4
enhanceosome; nuclear body; nuclear bodies that occur at super enhancers in mESCs
T
T
F
F
F
F
F
F
Buc BUC Danio rerio H0WFA5 Bucky ball
mitochondrial cloud; germ plasm; Balbiani body
T
T
F
T
F
T
T
F
Tdrd6 TDRD6 Danio rerio F1R237 Tudor domain-containing 6
mitochondrial cloud; germ plasm; Balbiani body
T
T
F
T
F
T
T
F
FtsZ FTSZ Escherichia coli P0A9A6 Cell division protein FtsZ
nuclear body; Ftsz-rich droplets at specific chromosomal DNA sites
F
T
F
T
F
F
T
T
SlmA SLMA Escherichia coli P0C093 Nucleoid occlusion factor SlmA
nuclear body; Ftsz-rich droplets at specific chromosomal DNA sites
F
T
F
T
F
F
T
T
Protein P P Rabies virus P22363 Phosphoprotein
cytoplasmic viral factory; Negri body
T
F
F
T
T
F
F
T
Protein N N Rabies virus Q8JXF6 Nucleoprotein
cytoplasmic viral factory; Negri body
T
F
F
T
T
F
F
T
RNase E RNE Caulobacter vibrioides Q9A749 Ribonuclease E
cytoplasmic ribonucleoprotein granule; bacterial RNP-bodies (BR-bodies)
T
T
F
F
T
F
F
F
Sam68 KHDRBS1 Homo sapiens Q07666 KH domain-containing, RNA-binding, signal transduction-associated protein 1
nuclear body; ribonucleoprotein complex; Sam68 nuclear bodies (SNBs)
T
F
F
T
T
T
F
T
HNRNPD HNRNPD Homo sapiens Q14103 Heterogeneous nuclear ribonucleoprotein D0
nuclear body; ribonucleoprotein complex; Sam68 nuclear bodies (SNBs)
T
F
F
T
T
T
F
T
SPOP SPOP Homo sapiens O43791 Speckle-type POZ protein
nuclear body; nuclear protein granule; SPOP/DAXX body
T
T
F
T
F
F
T
F
DAXX DAXX Homo sapiens Q9UER7 Death domain-associated protein 6
nuclear body; nuclear protein granule; SPOP/DAXX body
T
T
F
T
F
F
T
F
Oskar OSKA_DROME Drosophila melanogaster P25158-2 Isoform C of Maternal effect protein oskar
P granule
T
F
F
F
F
F
F
T
Histone H1 H11L_CHICK Gallus gallus P08287 Histone H1.11L
euchromatin
F
T
F
T
F
F
F
F
Sec16 SEC16 Drosophila melanogaster Q5BI67 LP14866p
Sec body; cytoplasmic protein granule
T
F
F
T
F
T
F
F
Sec24AB SEC24AB Drosophila melanogaster A1Z813 Sec24AB ortholog, isoform A
Sec body; cytoplasmic protein granule
T
F
F
T
F
T
F
F
Sec23 SEC23 Drosophila melanogaster Q9VNF8 Secretory 23, isoform A
Sec body; cytoplasmic protein granule
T
F
F
T
F
T
F
F
RPB1 POLR2A Homo sapiens P24928 DNA-directed RNA polymerase II subunit RPB1
RNA polymerase II, holoenzyme; POLII clusters
T
T
F
F
F
F
F
F
RPB1 RPO21 Saccharomyces cerevisiae P04050 DNA-directed RNA polymerase II subunit RPB1
RNA polymerase II, holoenzyme; POLII clusters
F
T
F
F
F
F
F
F
Cyclin-T1, CYCT1 CCNT1 Homo sapiens O60563 Cyclin-T1
nuclear speckle
T
T
F
F
F
F
F
F
DYRK1A DYRK1A Homo sapiens Q13627 Dual specificity tyrosine-phosphorylation-regulated kinase 1A
nuclear speckle
F
T
F
F
F
F
F
F
Protein P P Vesicular stomatitis Indiana virus P03520 Phosphoprotein
cytoplasmic viral factory; viroplasm
T
F
F
T
F
F
T
T
Protein N N Vesicular stomatitis Indiana virus P03521 Nucleoprotein
cytoplasmic viral factory; viroplasm
T
F
F
T
F
F
T
T
Protein L L Vesicular stomatitis Indiana virus P03523 RNA-directed RNA polymerase L
cytoplasmic viral factory; viroplasm
T
F
F
T
F
F
T
T
PrP PRNP Homo sapiens P04156 Major prion protein
cytoplasmic protein granule
T
T
F
T
F
F
T
F
Amyloid-beta oligomer APP Homo sapiens P05067 Amyloid-beta A4 protein
cytoplasmic protein granule
T
T
F
T
F
F
T
F
ADCP1 AT1G09320 Arabidopsis thaliana Q500V5 Agenet domain-containing protein
heterochromatin
T
T
F
T
F
T
T
F
CBX2 CBX2 Homo sapiens Q14781 Chromobox protein homolog 2
PcG protein complex; euchromatin; heterochromatin; PcG chromatin condensates
T
T
F
F
F
F
F
F
Std1 STD1 Saccharomyces cerevisiae Q02794 Protein STD1
cytoplasmic protein granule; Std1 bodies
T
F
F
T
F
F
F
F
Nab3 NAB3 Saccharomyces cerevisiae P38996 Nuclear polyadenylated RNA-binding protein 3
nuclear body
T
T
F
F
F
F
F
F
EPYC1 LCI5 Chlamydomonas reinhardtii Q94ET8 LCI5
pyrenoid
T
T
F
F
F
F
T
F
RIM1 RIMS1 Rattus norvegicus Q9JIR4 Regulating synaptic membrane exocytosis protein 1
cytoplasmic protein granule
F
T
F
T
F
F
T
F
RIM-BP2 RIMBP2 Rattus norvegicus Q9JIR1 RIMS-binding protein 2
cytoplasmic protein granule
F
T
F
T
F
F
T
F
VRN1 VRN1 Arabidopsis thaliana Q8L3W1 B3 domain-containing transcription factor VRN1
nuclear body
T
T
F
T
F
F
F
F
TIS11B ZFP36L1 Homo sapiens Q07352 mRNA decay activator protein ZFP36L1
cytoplasmic ribonucleoprotein granule; TIS granule
T
F
F
F
F
F
F
F
GATA3 GATA3 Homo sapiens P23771 Trans-acting T-cell-specific transcription factor GATA-3
enhanceosome; nuclear body; robustly E2-activated enhancers (MegaTrans) enhancers
T
T
F
F
F
F
F
F
ERα ESR1 Homo sapiens P03372 Estrogen receptor
enhanceosome; nuclear body; robustly E2-activated enhancers (MegaTrans) enhancers
T
T
F
F
F
F
F
F
DYRK3 DYRK3 Homo sapiens O43781 Dual specificity tyrosine-phosphorylation-regulated kinase 3
cytoplasmic stress granule; liquid-like DYRK3 speckles
T
F
F
F
F
F
F
F
SYN2 SYN2 Homo sapiens Q92777 Synapsin-2
presynaptic cytosol; a matrix holding together clusters of synaptic vesicles (SVs)
T
T
F
F
F
F
F
T
Cortactin CTTN Mus musculus Q60598 Src substrate cortactin
intracellular non-membrane-bounded organelle
F
T
F
F
F
F
F
F
Sup35 SUP35 Schizosaccharomyces pombe O74718 Eukaryotic peptide chain release factor GTP-binding subunit
cytoplasmic protein granule; Sup35 condensate
T
T
F
F
F
F
F
F
PML PML Homo sapiens P29590-12 Isoform PML-12 of Protein PML
PML body
T
F
F
T
F
F
T
T
PSD-95 DLG4_HUMAN Homo sapiens P78352-3 Isoform 3 of Disks large homolog 4
postsynaptic density
T
T
F
T
F
F
T
T
AGO2 AGO2 Homo sapiens Q9UKV8 Protein argonaute-2
RISC complex; P-body; micro-ribonucleoprotein complex; miRISC; GW-body
T
T
F
T
F
F
T
F
Rubisco RBCS-1 Chlamydomonas reinhardtii P00873 Ribulose bisphosphate carboxylase small chain 1, chloroplastic
pyrenoid
T
T
F
F
F
F
T
F
Mid1p MID1 Schizosaccharomyces pombe P78953 Division mal foutue 1 protein
contractile ring
T
T
F
F
F
T
F
F
Female sterile (1) Yb FS(1)YB Drosophila melanogaster Q9W4W2 Female sterile (1) Yb
Yb body
T
F
F
F
T
T
F
T
Matrin-3 MATR3 Homo sapiens P43243 Matrin-3
nuclear protein granule
T
F
F
F
F
F
F
F
HSP48 DDB_G0280215 Dictyostelium discoideum Q54VP4 Heat shock protein DDB_G0280215
intracellular non-membrane-bounded organelle
T
F
F
T
F
T
T
T
U2AF65 U2AF2 Homo sapiens P26368 Splicing factor U2AF 65 kDa subunit
nuclear speckle
T
T
F
F
F
F
T
F
MORC3 MORC3 Homo sapiens Q14149 MORC family CW-type zinc finger protein 3
MORC3-NBs
T
T
F
T
F
F
F
F
Rv1747 RV1747 Mycobacterium tuberculosis O65934 ABC transporter ATP-binding/permease protein Rv1747
intracellular non-membrane-bounded organelle
T
T
T
F
F
T
T
F
MSMEG_1642 MSMEG_1642 Mycobacterium smegmatis A0QSY1 ABC transporter, ATP-binding protein
intracellular non-membrane-bounded organelle
F
T
T
F
F
F
T
F
Nucleoprotein N Measles virus P35972 Nucleoprotein
inclusion body
T
F
F
T
F
F
T
F
YTHDF2, DF2 YTHDF2 Homo sapiens Q9Y5A9 YTH domain-containing family protein 2
P-body; cytoplasmic stress granule; neuronal ribonucleoprotein granule
T
T
F
F
F
F
F
F
YTHDF1, DF1 YTHDF1 Homo sapiens Q9BYJ9 YTH domain-containing family protein 1
P-body; cytoplasmic stress granule; neuronal ribonucleoprotein granule
T
T
F
F
F
F
F
F
YTHDF3, DF3 YTHDF3 Homo sapiens Q7Z739 YTH domain-containing family protein 3
P-body; cytoplasmic stress granule; neuronal ribonucleoprotein granule
T
T
F
F
F
F
F
F
TIAR-2 TIAR-2 Caenorhabditis elegans Q9U2F5 TIA-1/TIAL RNA binding protein homolog
axonal TIAR-2 granules
T
T
F
F
F
T
F
F
CPEB3 CPEB3 Homo sapiens Q8NE35 Cytoplasmic polyadenylation element-binding protein 3
P-body
T
T
F
T
T
T
F
F
Phosphoprotein P/V Measles virus P35974 Phosphoprotein
inclusion body
T
F
F
T
F
F
T
F
53BP1 TP53BP1 Homo sapiens Q12888 TP53-binding protein 1
nuclear body
T
T
F
F
F
F
F
T